The most likely cause is uneven development from too little chemistry in the Jobo tank, not simple agitation by itself. The repeated spacing matching the holder/tank support structure strongly suggests the film was being affected by the reel or holder geometry during processing. The abrupt horizontal line also points to solution level sitting at the same place repeatedly, which fits insufficient developer volume or incomplete coverage.

A less likely possibility is accidental fogging if the tank was opened during processing, with the reel structure projecting onto the film, but the consistent pattern across all sheets makes uneven chemical coverage the stronger explanation.

Best next step: run a controlled test with one sheet and verify the tank is loaded correctly and filled with the proper chemistry volume for rotary processing. If the marks repeat, the issue is in the tank setup/process rather than the film itself.